| University of Leeds | 617 | £200,360,912 |
| University of Oxford | 538 | £177,889,995 |
| University of Bristol | 509 | £151,121,812 |
| University of Edinburgh | 462 | £162,526,143 |
| University of Exeter | 435 | £147,164,220 |
| University of Southampton | 430 | £129,332,375 |
| University of Reading | 416 | £133,496,597 |
| University of Cambridge | 415 | £126,032,927 |
| Imperial College London | 395 | £154,036,183 |
| University of East Anglia | 389 | £109,333,000 |
| The University of Manchester | 361 | £117,111,676 |
| University College London | 361 | £116,485,348 |
| University of Liverpool | 282 | £84,368,525 |
| University of Sheffield | 273 | £78,213,102 |
| University of Birmingham | 272 | £90,337,004 |
| NERC CEH (Up to 30.11.2019) | 268 | £93,479,234 |
| University of York | 252 | £60,935,847 |
| NERC British Antarctic Survey | 244 | £97,810,987 |
| Newcastle University | 239 | £65,710,722 |
| Durham University | 237 | £65,406,721 |
| NOC (Up to 31.10.2019) | 234 | £73,123,058 |
| Lancaster University | 224 | £61,899,344 |
| University of Aberdeen | 220 | £55,870,879 |
| University of St Andrews | 197 | £52,739,323 |
| British Geological Survey | 190 | £39,674,595 |
| Cardiff University | 190 | £40,531,459 |
| University of Leicester | 164 | £48,036,573 |
| Plymouth Marine Laboratory | 161 | £48,633,618 |
| University of Glasgow | 154 | £40,807,293 |
| University of Plymouth | 151 | £29,494,536 |
| Bangor University | 139 | £37,315,493 |
| UK Centre for Ecology & Hydrology | 124 | £55,759,215 |
| National Oceanography Centre | 123 | £47,625,918 |
| The Natural History Museum | 117 | £19,769,893 |
| Scottish Association For Marine Science | 116 | £24,886,555 |
| The Open University | 106 | £15,551,574 |
| Queen Mary University of London | 99 | £20,611,338 |
| Swansea University | 95 | £21,152,160 |
| Royal Holloway, Univ of London | 94 | £17,717,756 |
| University of Stirling | 94 | £22,628,927 |
| University of Nottingham | 93 | £19,757,316 |
| University of Warwick | 91 | £23,166,741 |
| Cranfield University | 90 | £20,105,951 |
| King's College London | 85 | £23,719,413 |
| University of Hull | 77 | £12,640,392 |
| University of Essex | 71 | £15,736,079 |
| Aberystwyth University | 71 | £13,692,468 |
| University of Strathclyde | 67 | £29,773,402 |
| Heriot-Watt University | 67 | £26,510,999 |
| Loughborough University | 63 | £10,975,155 |
| Northumbria University | 60 | £14,551,360 |
| STFC - Laboratories | 59 | £12,112,088 |
| University of Bath | 54 | £15,221,155 |
| University of Sussex | 49 | £11,500,800 |
| Queen's University of Belfast | 41 | £5,256,488 |
| Marine Biological Association | 41 | £7,618,750 |
| University of Portsmouth | 33 | £6,948,884 |
| Zoological Soc London Inst of Zoology | 33 | £6,382,997 |
| University of Surrey | 31 | £7,104,063 |
| Rothamsted Research | 31 | £8,111,712 |
| Scottish Universities Env Research Cen | 28 | £4,561,885 |
| Manchester Metropolitan University | 25 | £6,319,779 |
| Keele University | 23 | £3,074,510 |
| University of Dundee | 23 | £4,323,023 |
| University of Hertfordshire | 22 | £3,133,132 |
| Royal Botanic Gardens Kew | 22 | £5,007,244 |
| University of Ulster | 21 | £2,336,349 |
| University of Kent | 21 | £2,764,084 |
| University of Bradford | 19 | £1,520,032 |
| Proudman Oceanographic Laboratory | 19 | £2,431,289 |
| Birkbeck College | 18 | £2,345,340 |
| Centre for Env Fisheries Aqua Sci CEFAS | 18 | £4,532,132 |
| University of Salford | 17 | £3,183,560 |
| University of Brighton | 17 | £2,965,968 |
| London School of Economics & Pol Sci | 16 | £3,635,884 |
| Bournemouth University | 16 | £1,576,806 |
| University of Lincoln | 16 | £3,678,186 |
| University of the West of England | 15 | £3,692,200 |
| The James Hutton Institute | 15 | £4,683,797 |
| Brunel University London | 13 | £2,454,139 |
| London Sch of Hygiene & Tropic. Medicine | 13 | £4,424,251 |
| Liverpool John Moores University | 12 | £1,429,987 |
| SRUC | 11 | £1,153,962 |
| SAHFOS | 11 | £628,068 |
| Royal Botanic Garden Edinburgh | 11 | £1,745,848 |
| Edinburgh Napier University | 10 | £1,670,469 |
| ODI | 10 | £1,629,703 |
| University of the Highlands and Islands | 8 | £977,418 |
| Royal Veterinary College | 8 | £1,535,442 |
| Roehampton University | 7 | £302,355 |
| Kingston University | 7 | £296,102 |
| Oxford Brookes University | 7 | £1,313,643 |
| British Trust for Ornithology | 6 | £625,872 |
| Institute of Development Studies | 6 | £3,390,621 |
| Met Office | 6 | £2,784,800 |
| University of Huddersfield | 6 | £914,783 |
| H R Wallingford Ltd | 6 | £1,085,256 |
| University of the West of Scotland | 5 | £1,300,926 |
| Harper Adams University | 5 | £407,254 |
| University of Gloucestershire | 5 | £778,554 |
| London and South East University Group | 5 | £1,155,131 |
| Diamond Light Source | 5 | £264,339 |
| Glasgow Caledonian University | 5 | £1,255,236 |
| Edge Hill University | 4 | £655,926 |
| Institute of Research for Dev (IRD) | 4 | £896,973 |
| RSPB | 4 | £558,860 |
| Robert Gordon University | 4 | £958,679 |
| City St George's, University of London | 4 | £1,176,369 |
| International Water Management Institute | 4 | £707,842 |
| Marine Scotland Science | 4 | £102,382 |
| Anglia Ruskin University | 3 | £730,782 |
| University of Worcester | 3 | £186,041 |
| Liverpool School of Tropical Medicine | 3 | £1,476,739 |
| University of Malawi | 3 | £509,827 |
| St George's University of London | 3 | £670,648 |
| NIAB | 3 | £646,549 |
| Fisheries Research Services Marine Labor | 3 | £476,482 |
| Abertay University | 3 | £639,927 |
| John Innes Centre | 3 | £886,797 |
| Fera Science Limited | 3 | £454,178 |
| Teesside University | 3 | £64,330 |
| National Museums of Scotland | 3 | £363,301 |
| University of Sunderland | 3 | £67,132 |
| CAB International | 2 | £340,646 |
| Moredun Research Institute | 2 | £504,673 |
| SEI Oxford Office Ltd | 2 | £248,949 |
| University of Derby | 2 | £128,664 |
| University of Lancashire | 2 | £284,741 |
| University of Wolverhampton | 2 | £543,773 |
| Conservation International Foundation | 2 | £885,095 |
| MetaMeta | 2 | £188,935 |
| Addis Ababa University | 2 | £301,669 |
| International Institute for Env and Dev | 2 | £409,883 |
| Coventry University | 2 | £258,451 |
| University of East London | 2 | £149,644 |
| Sokoine University of Agriculture | 2 | £349,684 |
| Bolivian Natura Foundation | 2 | £635,641 |
| The Alan Turing Institute | 2 | £514,302 |
| Nottingham Trent University | 2 | £241,672 |
| Inst of Grassland and Environmental Res | 2 | £195,803 |
| Wageningen University | 2 | £452,463 |
| De Montfort University | 2 | £59,508 |
| University of Westminster | 2 | £104,572 |
| CONDESAN | 2 | £322,467 |
| Basque Centre for Climate Change bc3 | 2 | £230,757 |
| Lilongwe Uni of Agri and Nat Resources | 2 | £152,220 |
| University of Cape Town | 2 | £2,028,130 |
| Makerere University | 2 | £130,538 |
| Falmouth University | 2 | £405,260 |
| University of Cumbria | 2 | £112,380 |
| Council for Sci and Industrial Res | 2 | £325,905 |
| SOAS University of London | 2 | £32,113 |
| Environment Agency | 2 | £285,528 |
| International Food Policy Research Inst | 2 | £608,731 |
| Glasgow School of Art | 2 | £25,035 |
| Flinders University of South Australia | 1 | £106,147 |
| Swedish Meteorological & Hydro Institute | 1 | £618,219 |
| Wellcome Sanger Institute | 1 | £153,088 |
| BBSRC Institute of Arable Crops Research | 1 | £28,105 |
| University of Milan-Bicocca | 1 | £146,700 |
| Ludwig Maximilian University of Munich | 1 | £248,906 |
| Leeds Beckett University | 1 | £4,923 |
| Carbon Foundation of East Africa | 1 | £87,597 |
| Institute of Occupational Medicine | 1 | £495,172 |
| IHE Delft Foundation | 1 | £1,931,581 |
| University of Dar es Salaam | 1 | £61,202 |
| Royal Northern College of Music | 1 | £4,923 |
| ICLEI - Local Govts for Sustainability | 1 | £211,597 |
| Medical Research Council (MRC) | 1 | £203,061 |
| WaterAid UK | 1 | £419,301 |
| University of Johannesburg | 1 | £177,065 |
| CNRS - Delegation Midi-Pyrenees | 1 | £172,281 |
| UK Health Security Agency | 1 | £303,816 |
| Policy Studies Institute | 1 | £11,954,596 |
| The Courtauld Institute of Art | 1 | £25,513 |
| Kenyatta University | 1 | £74,899 |
| University of Tasmania | 1 | £7,058 |
| Scottish Crop Research Institute | 1 | £2,531 |
| University of Nairobi | 1 | £221,661 |
| Stockholm University | 1 | £1,207,243 |
| Red Cross Red Crescent Climate Centre | 1 | £207,422 |
| Global Canopy Foundation | 1 | £152,599 |
| University of Geneva | 1 | £318,782 |
| Bangladesh Univ of Eng and Tech (BUET) | 1 | £795,644 |
| CORDIO East Africa | 1 | £9,119 |
| Intnl Union for Conservation of Nature | 1 | £47,753 |
| Stony Brook University | 1 | £285,924 |
| Hawassa University | 1 | £227,595 |
| Wildlife Conservation Society | 1 | £32,597 |
| National Physical Laboratory NPL | 1 | £1,945,909 |
| Nature Conservation Research Centre | 1 | £683,562 |
| The Francis Crick Institute | 1 | £76,538 |
| World Agroforestry Centre | 1 | £146,827 |
| Rhodes University | 1 | £62,127 |
| University of London | 1 | £5,423 |
| Macaulay Institute | 1 | £64,390 |
| World Conservation Monitoring Cen WCMC | 1 | £120,496 |
| Bath Spa University | 1 | £7,731 |
| BBSRC John Innes Centre | 1 | £297,676 |
| Birmingham City University | 1 | £27,059 |
| University of Chester | 1 | £20,012 |
| Public Health England | 1 | £439,818 |
| University of Bedfordshire | 1 | £4,923 |
| Macaulay Land Use Research Institute | 1 | £152,911 |
| University of California, Santa Cruz | 1 | £7,182 |
| Indian Institute of Technology Roorkee | 1 | £242,198 |
| Aston University | 1 | £265,781 |
| START International Inc | 1 | £256,406 |
| University of Sao Paulo | 1 | £12,106 |
| University of South Wales | 1 | £138,232 |
| Int Centre for Tropical Agriculture | 1 | £778,003 |
| University of Maiduguri | 1 | £15,960 |
| Bradford Teaching Hosp NHS Found Trust | 1 | £370,805 |
| EMBL - European Bioinformatics Institute | 1 | £43,274 |
| North Carolina State University | 1 | £181,569 |
| Southern Agricultural Research Institute | 1 | £315,311 |
| UK Astronomy Technology Centre | 1 | £18,831 |
| CIRAD | 1 | £151,874 |
| ILRI (International Livestock Res Inst) | 1 | £471,287 |
| Geological Survey of Ethiopia | 1 | £22,168 |
| Royal Central Sch of Speech and Drama | 1 | £5,423 |
| Kulima Integrated Development Solutions | 1 | £576,346 |
| Goldsmiths College | 1 | £119,059 |
| National Nuclear Laboratory (NNL) | 1 | £309,181 |
| Zutari | 1 | £183,349 |
| Cardiff Metropolitan University | 1 | £7,731 |
| Inst for European Environmental Policy | 1 | £162,166 |
| Laboratory of Radioisotopes | 1 | £181,031 |
| Institute of Urban Environment | 1 | £26,636 |
| University of Central Asia | 1 | £227,225 |
| University of Glamorgan | 1 | £107,726 |
| International Potato Centre | 1 | £51,996 |
| Royal College of Art | 1 | £21,261 |
| The British Museum | 1 | £6,008 |
| University of Bonn | 1 | £141,779 |
| The National Trust | 1 | £153,021 |
| East China Normal University | 1 | £14,954 |
| York St John University | 1 | £45,657 |
| Sheffield Hallam University | 1 | £11,597 |
| Catholic University of Peru (PUCP) | 1 | £18,750 |
| University of KwaZulu-Natal | 1 | £208,765 |
| Middlesex University | 1 | £149,162 |
| CSIRO | 1 | £14,108 |
| University of the Arts London | 1 | £34,017 |
| Weber State University | 1 | £93,720 |
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